Excerpt from Quarterly Journal of Microscopical Science, 1921, Vol. 65
The first suggestion of any structure at all observable in chromosomes seems to be due to Pfitzner ('Morph. Jahrb.' , vii, 1882), who suggested that a chromosome is made up of a row of granules of chromatin embedded in an achromatic or less chromatic thread. Belief in these granules - later dignified by the names of 'chromomeres,' 'chromioles,' and the like - long held sway, and still lingers in many minds. I do not think it necessary to enter into a detailed discussion of this view; for I think it is now indubitable that the supposed granules are nothing but the misinterpreted images of twists of the chromosome, or of bulges in it. The figures illustrating this paper afford abundant instances of bulges caused by twists of the chromosomes; and those illustrating my paper on the chromosomes of Paris quadrifo1ia('La Cellule,' xxviii, 2, 1912, p. 265) of bulges caused by alveoles in them; either of which, if indistinctly seen, may lend themselves to an erroneous interpretation as granules.
At the present time two other theories are in the field: the chromonema theory, and the alveolation theory.
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